The Lion of St. Mark is a representation of the evangelist St. Mark (and is also the symbol of Venice). The winged beast is holding the book of St. Mark. The mythology says that a lion is one of four animals that hangs out around the throne of God -- because of course God likes cats -- but beyond that there is little agreement on the symbolism.
All coin images in Daniel's Coin Zoo are from my personal collection. I collect, research, and personally photograph every coin displayed on this site. PLEASE do not take my images without permission. If you would like to use any coin image you see, just ask me. Thank you.